In these 2 studies, other species of fish were tested to see if they could be carriers of KHV. 

While several species were susceptible to KHV and some died from the exposure, neither study could conclusively prove that the other species infected with KHV could then infect naive Koi.  Read the studies here:

CLICK HERE for the study with Gudgeon.

CLICK HERE for the study with Stone Loach, and Sterlet/Beluga hybrids.